553 error when sending mail (self-test included) after setting up monitored inbox

Mautic 4.4.7
My PHP version is : 7.4.33
MariaDB version is 10.5.18-MariaDB-0+deb11u1

Updating/Installing Errors none

Upgrading/installing via (delete as applicable) : Command Line

Everything works fine until I set up monitored inbox settings; What’s strange it used to work fine, but now it isn’t

Case #1

  1. Other SMTP sserver:

Email address to send mail from: zzz@mysite.online

Username for the selected mail service: zzz@mysite.online

SMTP Port 465

SSL

Test connection _> OK

Ssend test email → OK

Case #2

  1. Other SMTP sserver:

Email address to send mail from: zzz@mysite.online

Username for the selected mail service: zzz@mysite.online

SMTP Port 465

SSL

Monitored address : zzz@mysite.online

IMAP username: zzz@mysite.online

Port 993

SSL

IMAP server xxx

Test connection and fetch folders→ OK

Test connection → OK

Send test email: Expected response code 250 but got code “553”, with message "553 5.7.1 Sender address rejected: not owned by auth user

Mautic’s log

I have no idea what’s going on and why it has started happening.

Any idea how to fix it?

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